David W. Greenfield, Terry Greenfield, Introgressive Hybridization between Gila orcutti and Hesperoleucus symmetricus (Pisces: Cyprinidae) in the Cuyama River Basin, California: I. Meristics, Morphometrics and Breeding, Copeia, Vol. 1972, No. 4 (Dec. 29, 1972), pp. 849-859
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Greenfield et al. (1972) studied this question.
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