C. Alana Tibbets, Thomas E. Dowling, Effects of Intrinsic and Extrinsic Factors on Population Fragmentation in Three Species of North American Minnows (Teleostei: Cyprinidae), Evolution, Vol. 50, No. 3 (Jun., 1996), pp. 1280-1292
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Tibbets et al. (1996) studied this question.
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