Jukka Suhonen, Rauno V. Alatalo, Allan Carlson, , Food Resource Distribution and the Organization of the Parus Guild in a Spruce Forest, Ornis Scandinavica (Scandinavian Journal of Ornithology), Vol. 23, No. 4 (Oct. - Dec., 1992), pp. 467-474
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Suhonen et al. (1992) studied this question.
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