Laboratory measurements indicate that observed differences in the rate of water‐loss from inactive Helix aspersa, Otala lactea and Sphincterochila boissieri are due to morphological adaptations such as a thicker shell, reduced aperture and thicker epiphragm. These modifications enable terrestrial snails to inhabit dryer environments by increasing the length of time the snail can remain inactive.
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J. Machin (1967) studied this question.
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