Noriaki Murakami, Satoru Nogami, Mikio Watanabe, Kunio Iwatsuki, Phylogeny of Aspleniaceae Inferred from rbcL Nucleotide Sequences, American Fern Journal, Vol. 89, No. 4 (Oct. - Dec., 1999), pp. 232-243
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Murakami et al. (1999) studied this question.
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