Giuseppe Nascetti, Roberta Cimmaruta, Benedetto Lanza, Luciano Bullini, Molecular Taxonomy of European Plethodontid Salamanders (Genus Hydromantes), Journal of Herpetology, Vol. 30, No. 2 (Jun., 1996), pp. 161-183
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Nascetti et al. (1996) studied this question.
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