Michael A. Steele, Travis Knowles, Kenneth Bridle, Ellen L. Simms, Tannins and Partial Consumption of Acorns: Implications for Dispersal of Oaks by Seed Predators, The American Midland Naturalist, Vol. 130, No. 2 (Oct., 1993), pp. 229-238
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Steele et al. (1993) studied this question.
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