J. C. Coulson, J. C. Horobin, Jennifer Butterfield, G. R. J. Smith, The Maintenance of Annual Life-Cycles in two Species of Tipulidae (Diptera); A Field Study Relating Development, Temperature and Altitude, Journal of Animal Ecology, Vol. 45, No. 1 (Feb., 1976), pp. 215-233
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Coulson et al. (1976) studied this question.
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