J. Bastow Wilson, Habiba Gitay, Stephen H. Roxburgh, Warren McG. King, Raymond S. Tangney, Egler's Concept of 'Initial Floristic Composition' in Succession: Ecologists Citing It Don't Agree What It Means, Oikos, Vol. 64, No. 3 (Sep., 1992), pp. 591-593
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Wilson et al. (1992) studied this question.
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