Daniel Perea, , , First Fossil Record (Late Miocene) of Phrynops (P. geoffroanus Complex: Chelidae) from Uruguay: Biostratigraphical and Paleoenvironmental Context, Copeia, Vol. 1996, No. 2 (May 16, 1996), pp. 445-451
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Perea et al. (1996) studied this question.
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