D. S. Ranwell, E. C. F. Bird, J. C. E. Hubbard, R. E. Stebbings, Spartina Salt Marshes in Southern England: V. Tidal Submergence and Chlorinity in Poole Harbour, Journal of Ecology, Vol. 52, No. 3 (Nov., 1964), pp. 627-641
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Ranwell et al. (1964) studied this question.
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