D. R. Livingstone, P. Garcia Martinez, X. Michel, J. F. Narbonne, S. O'Hara, D. Ribera, G. W. Winston, Oxyradical Production as a Pollution-Mediated Mechanism of Toxicity in the Common Mussel, Mytilus edulis L., and Other Molluscs, Functional Ecology, Vol. 4, No. 3, New Horizons in Ecotoxicology (1990), pp. 415-424
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Livingstone et al. (1990) studied this question.