Javier Francisco-Ortega, Leslie R. Goertzen, Arnoldo Santos-Guerra, Abdelmalek Benabid, Robert K. Jansen, Molecular Systematics of the Asteriscus Alliance (Asteraceae: Inuleae) I: Evidence from the Internal Transcribed Spacers of Nuclear Ribosomal DNA, Systematic Botany, Vol. 24, No. 2 (Apr. - Jun., 1999), pp. 249-266
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Francisco‐Ortega et al. (1999) studied this question.
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