Marta M. Cestari, Pedro M. Galetti, Jr., Chromosome Studies of Serrasalmus spilopleura (Characidae, Serrasalminae) from the Paraná-Paraguay Rivers: Evolutionary and Cytotaxonomic Considerations, Copeia, Vol. 1992, No. 1 (Feb. 3, 1992), pp. 108-112
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Cestari et al. (1992) studied this question.
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