Nikolai Lebedev, Barbara van Cleve, Gregory Armstrong, Klaus Apel, Chlorophyll Synthesis in a Deetiolated (det340) Mutant of Arabidopsis without NADPH-Protochlorophyllide (PChlide) Oxidoreductase (POR) A and Photoactive PChlide-F655, The Plant Cell, Vol. 7, No. 12 (Dec., 1995), pp. 2081-2090
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Lebedev et al. (1995) studied this question.