Gregory A. Armstrong, Bernd Weisshaar, Klaus Hahlbrock, Homodimeric and Heterodimeric Leucine Zipper Proteins and Nuclear Factors from Parsley Recognize Diverse Promoter Elements with ACGT Cores, The Plant Cell, Vol. 4, No. 5 (May, 1992), pp. 525-537
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Armstrong et al. (1992) studied this question.