C. R. Eskey, V. H. Haas, Plague in the Western Part of the United States: Infection in Rodents, Experimental Transmission by Fleas, and Inoculation Tests for Infection, Public Health Reports (1896-1970), Vol. 54, No. 32 (Aug. 11, 1939), pp. 1467-1481
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Eskey et al. (1939) studied this question.