Kathleen G. Dwyer, Muthugapatti K. Kandasamy, Dusty I. Mahosky, Joann Acciai, Bela I. Kudish, Janet E. Miller, Mikhail E. Nasrallah, June B. Nasrallah, A Superfamily of S Locus-Related Sequences in Arabidopsis: Diverse Structures and Expression Patterns, The Plant Cell, Vol. 6, No. 12 (Dec., 1994), pp. 1829-1843
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Dwyer et al. (1994) studied this question.