Ian D. Hodkinson, N. R. Webb, J. S. Bale, W. Block, S. J. Coulson, A. T. Strathdee, Global Change and Arctic Ecosystems: Conclusions and Predictions from Experiments with Terrestrial Invertebrates on Spitsbergen, Arctic and Alpine Research, Vol. 30, No. 3 (Aug., 1998), pp. 306-313
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Hodkinson et al. (1998) studied this question.
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