John A. S. Nagy, Mark A. Haroldson, Comparisons of Some Home Range and Population Parameters among Four Grizzly Bear Populations in Canada, Bears: Their Biology and Management, Vol. 8, A Selection of Papers from the Eighth International Conference on Bear Research and Management, Victoria, British Columbia, Canada, February 1989 (1990), pp. 227-235
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Nagy et al. (1990) studied this question.
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