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A relationship between yeast flocculation and intracellular acid-soluble glycogen has been established which has been substantiated using flocculation mutants (mutants with altered capacities to flocculate) as well as a normal strain of Saccharomyces carlsbergensis. Sound evidence exists to implicate physiological differences in carbohydrate metabolism (glycogen storage) to this physical property of brewing significance.
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Patel et al. (1975) studied this question.
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