Yutaka Watanuki, Akiko Kato, Yoshihisa Mori, Yasuhiko Naito, Diving Performance of Adélie Penguins in Relation to Food Availability in Fast Sea-Ice Areas: Comparison between Years, Journal of Animal Ecology, Vol. 62, No. 4 (Oct., 1993), pp. 634-646
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Watanuki et al. (1993) studied this question.
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