A modification of the original Helm et al. method was used to determine the flocculation of a moderately flocculent strain of Saccharomyces carlsbergensis. This test appeared to be more useful than methods adapted from Greenshields or Kato & Nishikawa, and made it possible to monitor flocculation trends during the course of fermentation. The effects of calcium, pH, ethanol, wort composition and yeast physiological age on the flocculation of Saccharomyces carlsbergensis were also studied.
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Patel et al. (1975) studied this question.
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