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H. F. Lamb, U. Eicher, V. R. Switsur, An 18,000-Year Record of Vegetation, Lake-Level and Climatic Change from Tigalmamine, Middle Atlas, Morocco, Journal of Biogeography, Vol. 16, No. 1 (Jan., 1989), pp. 65-74
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Lamb et al. (1989) studied this question.
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