Russel A. Kramm, Kenneth R. Kramm, Activities of Certain Species of Eleodes in Relation to Season, Temperature, and Time of Day at Joshua Tree National Monument (Coleoptera: Tenebrionidae), The Southwestern Naturalist, Vol. 16, No. 3/4 (Feb. 18, 1972), pp. 341-355
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Kramm et al. (1972) studied this question.
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