Odd Terje Sandlund, Karl Gunnarsson, , Bror Jonsson, Torfinn Lindem, , Hilmar J. Malmquist, , , Sigurdur S. Snorrason, The Arctic Charr Salvelinus alpinus in Thingvallavatn, Oikos, Vol. 64, No. 1/2, Ecology of Oligotrophic, Subarctic Thingvallavatn (May - Jun., 1992), pp. 305-351
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Sandlund et al. (1992) studied this question.
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