H. C. J. Godfray, J. K. Waage, Predictive Modelling in Biological Control: The Mango Mealy Bug (Rastrococcus invadens) and Its Parasitoids, Journal of Applied Ecology, Vol. 28, No. 2 (Aug., 1991), pp. 434-453
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Godfray et al. (1991) studied this question.
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