In a study based on morphological, embryological and chemical data of thc Plantaginaceae, within the subclass Sympetalae or Asteridae, the superorder 1,amianae is shown to be monophyletic. However, it was not possible to reconstruct the phylogeny within Lamianae or to find a sister-,group for the monophyletic Plantaginaceae -Hydrostachyaceae for the latter is rejected. Thrcr or rarely four genera have previously been recognized within Plantaginaceae, but in both cases Plantago appears as polyphyletic, which is considered unacceptable. Six clades are recognized as subgenera within Plantago:
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Knud Rahn (1996) studied this question.
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