Peter J. Grubb, Daniel J. Metcalfe, E. A. Anne Grubb, Glyn D. Jones, Nitrogen-Richness and Protection of Seeds in Australian Tropical Rainforest: A Test of Plant Defence Theory, Oikos, Vol. 82, No. 3 (Sep., 1998), pp. 467-482
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Grubb et al. (1998) studied this question.
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