Randall L. Morrison, Matthew S. Rand, Sally K. Frost-Mason, Cellular Basis of Color Differences in Three Morphs of the Lizard Sceloporus undulatus erythrocheilus, Copeia, Vol. 1995, No. 2 (May 3, 1995), pp. 397-408
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Morrison et al. (1995) studied this question.
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