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A generalized FORTRAN IV computer program (NEGST) for nested gene diversity analysis and its application to brown trout gene frequency data are presented. In its present shape the program handles up to five hierarchical levels of subdivision, 70 operational units (subpopulations), and 50 polymorphic loci with 20 alleles per locus.
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Chakraborty et al. (2008) studied this question.
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