John O. Corliss, D. Wayne Coats, A New Cuticular Cyst-Producing Tetrahymenid Ciliate, Lambornella clarki n. sp., and the Current Status of Ciliatosis in Culicine Mosquitoes, Transactions of the American Microscopical Society, Vol. 95, No. 4 (Oct., 1976), pp. 725-739
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Corliss et al. (1976) studied this question.
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