In the cockroach there are two distinct sets of photoreceptors-the compound eyes and the ocelli-which may function as a sensory input whereby circadian locomotor rhythms are entrained by environmental light cycles. Surgical removal of the ocelli did not interfere with normal entrainment, but covering over the compound eyes effected a loss of entrainment.
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Shepherd K. de F. Roberts (1965) studied this question.
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