Karen C. Cone, Suzy M. Cocciolone, Cynthia A. Moehlenkamp, Timothy Weber, Bruce J. Drummond, Laura A. Tagliani, Benjamin A. Bowen, Gary H. Perrot, Role of the Regulatory Gene pl in the Photocontrol of Maize Anthocyanin Pigmentation, The Plant Cell, Vol. 5, No. 12 (Dec., 1993), pp. 1807-1816
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Cone et al. (1993) studied this question.