Himadri B. Pakrasi, Bruce A. Diner, John G. K. Williams, Charles J. Arntzen, Deletion Mutagenesis of the Cytochrome b₅₅₉ Protein Inactivates the Reaction Center of Photosystem II, The Plant Cell, Vol. 1, No. 6 (Jun., 1989), pp. 591-597
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Pakrasi et al. (1989) studied this question.