Seedlings of most of the Gramineae possess a mesocotyl, but a few tribes are characterized by its absence. Seedlings of Bromus resemble the Triticeae in having no mesocotyl and this is further evidence for separating the Bromeae both from the Festuceae and from the Brachypodiae.
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D. J. Harberd (1972) studied this question.
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