John Roberts, Christine F. Pymar, J. S. Wallace, Rona M. Pitman, Seasonal Changes in Leaf Area, Stomatal and Canopy Conductances and Transpiration from Bracken Below a Forest Canopy, Journal of Applied Ecology, Vol. 17, No. 2 (Aug., 1980), pp. 409-422
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Roberts et al. (1980) studied this question.
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