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Mark R. Alfenito, Erik Souer, Christopher D. Goodman, Robin Buell, Jos Mol, Ronald Koes, Virginia Walbot, Functional Complementation of Anthocyanin Sequestration in the Vacuole by Widely Divergent Glutathione S-Transferases, The Plant Cell, Vol. 10, No. 7 (Jul., 1998), pp. 1135-1149
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Alfenito et al. (1998) studied this question.
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