Estimating Attack and Brood Totals and Densities of the Mountain Pine Beetle in Individual Lodgepole Pine Trees

A method was developed for estimating attack and brood totals of the mountain pine beetle in part or all of the infested bole of individual lodgepole pine trees. The method requires measurement of tree height, tree diameter at 1.37 m, and maximum height of attack, and tallying of brood and attacks on paired 20 cm by 30 cm samples centered at 1.22 m on the bole. Attack and brood totals are estimated in two ways: (1) as a product of the variables ( a ) total bole surface area, ( b ) proportion of bole surface area infested, and ( c ) mean attack/brood density; and (2) regression on the combined variables of tree diameter, infested height, and attack/brood density at 1.22 m on the bole. An approach to developing variance estimates for brood and attack totals is discussed.