Eileen V. Carey, Sandra Brown, Andrew J. R. Gillespie, Ariel E. Lugo, Tree Mortality in Mature Lowland Tropical Moist and Tropical Lower Montane Moist Forests of Venezuela, Biotropica, Vol. 26, No. 3 (Sep., 1994), pp. 255-265
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Carey et al. (1994) studied this question.
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