Robert W. Harrington, Jr., How Ecological and Genetic Factors Interact to Determine When Self-Fertilizing Hermaphrodites of Rivulus marmoratus Change into Functional Secondary Males, with a Reappraisal of the Modes of Intersexuality among Fishes, Copeia, Vol. 1971, No. 3 (Sep. 3, 1971), pp. 389-432
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Robert W. Harrington (1971) studied this question.
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