J. Grace, H. W. Woolhouse, A Physiological and Mathematical Study of the Growth and Productivity of a Calluna-sphagnum Community. II. Light Interception and Photosynthesis in Calluna, Journal of Applied Ecology, Vol. 10, No. 1 (Apr., 1973), pp. 63-76
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Grace et al. (1973) studied this question.
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