E. E. Sandison, M. B. Hill, The Distribution of Balanus pallidus stutsburi Darwin, Gryphaea gasar ((Adanson) Dautzenberg), Mercierella enigmatica Fauvel and Hydroides uncinata (Philippi) in Relation to Salinity in Lagos Harbour and Adjacent Creeks, Journal of Animal Ecology, Vol. 35, No. 1 (Feb., 1966), pp. 235-250
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Sandison et al. (1966) studied this question.