The movements of some of the Medusae (e. g. Sarsia) appear to be as voluntary as are those of insects. Some of the discophorous species of naked-eyed Medusae*, when threatened with injury, manifest peculiar movements, which are quite distinct from the ordinary locomotor contractions. These movements consist in a very strong and protracted systole, followed by a slow and gradual diastole. This spasm-like series of movements is never performed by any Medusa except when the animal is being injured or threatened with injury.
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George John Romanes (1876) studied this question.