P. Laan, M. J. Berrevoets, S. Lythe, W. Armstrong, C. W. P. M. Blom, Root Morphology and Aerenchyma Formation as Indicators of the Flood-Tolerance of Rumex Species, Journal of Ecology, Vol. 77, No. 3 (Sep., 1989), pp. 693-703
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Laan et al. (1989) studied this question.
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