Among the 56 beneficial insects collected and observed associated with pear psylla in France, 16 were abundant ; these arthropods included 2 parasitoids, Prionomitus mitratus (Dalman) and Trechnites psyllae (Ruschka) (Encyrtidae), and 14 predators, Anthocoris nemoralis (F.), A. nemorum (L.), Orius horvathi (Reuter), O. vicinus (Ribaut) (Het. : Anthocoridae) ; Heterotoma meriopterum (Scopoli), Campyloneura virgula (Herrich-Schaffer), Orthotylus nassatus (F.) (Het. : Miridae) ; Stethorus punctillum (Weise), Scymnus rubromaculatus (Goeze), S. subvillosus (Goeze), Coccinella septempunctata L. (Col. Coccinellidae) ; Chrysoperla carnea (Stephens) (Neur : Chrysopidae) ; Coniopteryx borealis Tjeder (Neur. : Coniopterygidae) and the mite Allothrombium fuliginosum Hermann. All these species are polyphagous but only the first three are tied to pear psylla. It was shown that the environment of the orchards, comprising hawthorn and nettle, is a reservoir for the main pear psylla enemies. Other prey or hosts and host plants of these beneficial arthropods are cited from recent literature.
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Franck Hérard (1986) studied this question.