This study examines the within- and between-population variation in components of yield in R. chamaemorus. Specifically, it addresses the following questions: are there any consistent differences in seed size or total allocation to seeds between populations of open and shaded habitats? Is seed size negatively correlated with yield components like seed number, number of seeds per unit leaf mass, or number of seeds per unit rhizome mass? How do supplemental pollination and defoliation affect seed number and size?
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Jon Ågren (1989) studied this question.