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Ronelle Roth, Lisa N. Hall, Thomas P. Brutnell, Jane A. Langdale, bundle sheath defective2, a Mutation That Disrupts the Coordinated Development of Bundle Sheath and Mesophyll Cells in the Maize Leaf, The Plant Cell, Vol. 8, No. 5 (May, 1996), pp. 915-927
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Roth et al. (1996) studied this question.
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