Triple-strand beta-sheets having up- and -down topology are widespread in proteins and occur in two forms denoted here as S-like and Z-like beta-sheets. In many cases they are included in super-secondary structures of higher order. A number of such structures is described in this paper. An important feature of these super-secondary structures is that they have a unique handedness. Another feature is that some of them only involve S-like beta-sheets and others only Z-like beta-sheets.
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А. В. Ефимов (1993) studied this question.
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