Manuel B. Sainz, Erich Grotewold, Vicki L. Chandler, Evidence for Direct Activation of an Anthocyanin Promoter by the Maize C1 Protein and Comparison of DNA Binding by Related Myb Domain Proteins, The Plant Cell, Vol. 9, No. 4 (Apr., 1997), pp. 611-625
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Sainz et al. (1997) studied this question.